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There is often a close relation to the portal tract, although more scattered groups of proliferating oval cells can be seen diffusely throughout the liver following xenobiotic-induced hepatic injury (Engelhardt 1997)

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For example, cancer cells produce lactate through glycolysis to satisfy their energy needs even in the presence of oxygen, known as the Warburg effect (Ward and Thompson, 2012)

Its copper binding property is particularly important, as it helps limit metal catalyzed oxidative stress, thereby protecting cells from extensive damage and influencing downstream signaling pathways