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It was first isolated and characterized in 1993 by researchers at the University of Zagreb in Croatia, who derived it from a larger protein naturally found in human gastric juice

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Cells were then washed three times with ice-cold PBS and incubated for 1 h at 4 C with 1 mM bis(sulfosuccinimidyl) suberate (BS3, non-cell permeable cross-linker, Thermo Scientific) in PBS

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